Melitaea didyma (Esper, 1778) NW.China. See [maps] Red-Band Fritillary Roter Scheckenfalter Spotted Fritillary Doncella timida Przeplatka didyma

• Papilio didyma Esper, 1778; Die Schmett. Th. I, Bd. 1 (7): pl. 41, f. 3
• ?Papilio didyma Esper, 1781; Die Schmett. Th. I, Bd. 2 (2): pl. 61, f. 1; TL: Bavaria
• Papilio cytheris Mueschen, 1781
• Papilio athulia Fabricius, 1787; Mantissa Insectorum 2: 59
• Melitaea didyma v. dalmatina Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.)
• dalmatina Staudinger, 1870
• didyma ab. nominoe Oberthür, 1900; Bull. Soc. ent. Fr. 1900: 276, pl. 1 [?? not named]
• ?Melitaea didyma ab. nominoe Oberthür, 1909; Étud. Lépid. Comp. 3: 411, pl. 26, f. 136
• Militaea[sic] didyma ab. ziegleri Stichel, 1900; Berl. ent. Z. 45: 128, pl. 2, f. 5
• ?jaika Krulikowsky, 1897; Mat. pozn. F.F. Ross. Imp. 3: 322 [?]
• jaika Krulikowsky, 1901; Mat. k. pozn. F.F. Ross. Imp. 6: 31
• Melitaea didyma ab. pudica Stichel, 1901; Berl. ent. Z. 46: 23
• Melitaea didyma ab. livida Klemensiewicz, 1906; Verh. zool.-bot. ges. Wien 56: 160
• Melitaea didyma ab. kempeleni Aigner-Abafi, 1906; Rovart. Lapok 13 (4): 71
• Melitaea didyma ab. kempeleni Aigner-Abafi, 1906; Ent. Zs. 19 (35): 208; TL: Mátra-Gebirge
• Melitaea didyma ab. fischeri Aigner-Abafi, 1906; Rovart. Lapok 13 (4): 72
• Melitaea didyma ab. fischeri Aigner-Abafi, 1906; Ent. Zs. 19 (35): 208; TL: Kassa
• Melitaea didyma ab. radiata Aigner-Abafi, 1906; Ann. hist.-nat. Mus. Hung. 4 (2): 494
• Melitaea didyma ab. striata Skala, 1907; Ent. Zs. 20 (42): 310, pl. 4, f. 1
• Melitaea didyma ab. nigra Skala, 1907; Ent. Zs. 20 (42): 310, pl. 4, f. 2
• Melitaea didyma ab. fasciata Skala, 1907; Ent. Zs. 20 (42): 310, pl. 4, f. 3
• Melitaea didyma ab. marginata Skala, 1907; Ent. Zs. 20 (42): 310, pl. 4, f. 4
• Melitaea didyma ab. tenuisignata Skala, 1907; Ent. Zs. 20 (42): 310, pl. 4, f. 5
• Melitaea didyma ab. ocellata Skala, 1907; Ent. Zs. 20 (42): 311, pl. 4, f. 6
• Melitaea didyma ab. interrupta Skala, 1907; Ent. Zs. 20 (42): 311, pl. 4, f. 7
• Melitaea didyma ab. zinburgi Skala, 1907; Ent. Zs. 20 (42): 311, pl. 4, f. 9
• Melitaea didyma ab. pallida Skala, 1907; Ent. Zs. 20 (42): 311, pl. 4, f. 10
• Melitaea didyma ab. oblongomaculata Skala, 1908; Ent. Zs. 21: 238
• Melitaea didyma ab. ochracea Skala, 1908; Ent. Zs. 21: 238
• Melitaea didyma ab. nigrita Skala, 1908; Ent. Zs. 21: 268 (repl. Melitaea didyma ab. nigra Skala, 1907)
• Melitaea didyma ab. ♀ nigerrima Schultz, 1908; Soc. Ent. 22 (23): 177
• Melitaea didyma ab. ♀ speciosa Schultz, 1908; Ent. Zs. 22 (23): 91
• pieszczeki Skala, 1909; Ent. Jahrb. 19: 136
• Melitaea didyma ab. albicans Pieszczek, 1910; Jber. Wien. Ent. Ver. 20: 85
• Melitaea didyma ab. radiata Oberthür, 1909; Étud. Lépid. Comp. 3: 411, pl. 26, f. 137, 140, 143
• Melitaea didyma ab. wullschlegeli Oberthür, 1909; Étud. Lépid. Comp. 3: 411, pl. 26, f. 141-142, 145
• progressiva Ksienschopolski, 1911; Trudy obshch.Izsl. Volyni Zitomir 8: 37, pl. 2, f. 7
• minor Ksienschopolski, 1911; Trudy obshch.Izsl. Volyni Zitomir 8: 37
• striata Ksienschopolski, 1911; Trudy obshch.Izsl. Volyni Zitomir 8: 38, pl. 2, f. 8
• Melitaea didyma ab. emicandida Verity, 1911; Boll. Soc. ent. Ital. 42: 266, pl. 1, f. 2
• acrogynoides Reverdin, 1912; Bull. Soc. Lép. Genève 2 (1): (41-50)
• anticoradiata de Saussure, 1914; Bull. Soc. Lép. Genève 3: 78, pl. 1, f. 9
• mauritii Reverdin, 1918; Bull. Soc. Lep. Genève 4: 40
• Melitaea didyma (I) gen. rubida Verity, 1919; Ent. Rec. J. Var. 31: 179
• Melitaea didyma r. pumila Verity, 1919; Ent. Rec. J. Var. 31: 181
• Melitaea didyma protea (II) gen. caldaria Verity, 1920; Boll. Lab. Zool. Gen. Agr. Portici. 14: 60
• Melitaea didyma gen. autumn. ab. latonigenoides Kesenheimer, 1921; Ent. Rundsch. 38: 38
• Melitaea didyma (II) gen. romula Verity, 1922; Ent. Rec. J. Var. 34: 12
• Melitaea didyma ab. marginimaculata Kautz, 1923; Verh. zool.-bot. Ges. Wien 69 (s.b.): 123
• mireio Foulquier, 1923; Cat. Lep. France: 60
• bockletii Heuer, 1924; Int. ent. Zs. 17: 168
• depauperata de Sagarra, 1924; Trab. mus. Ciénc. nat. Barcelona 11 (1): 14
• moulinsi Trimoulet, 1924; Bull. Soc. Linn. Bordaeux 22: 17
• nigroguttata Sovinsky, 1927; Trav. Sta. biol. Dniepre 2: 381 (repl. striata Ksienschopolski, 1911)
• Melitaea didyma ab. majellensis Dannehl, 1927; Mitt. Münch. Ent. Ges. 17 (1-6): 2
• Melitaea didyma ab. kalugini Kardakoff, 1928; Ent. Mitt. 17 (4): 270
• Melitaea didyma ab. robiginosa Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 3
• Melitaea didyma gen. ignea Verity, 1929; Ent. Rec. J. Var. 41 (5): 74; TL: Intra
• Melitaea didyma ab. radiata Herold, 1930; Carinthia II 39-40: 65
• nigralba Febvay du Couedic, 1934; Lambillionea 34: 150
• cumulans Skala, 1936; Arb. ent. Abt. Landesmus Brünn, Nr. 3: 16
• storacei (Berio, 1942) (Melitaeta); Boll. Soc. Ent. Ital. 74: 100
• unifasciata Caruel, 1944; Misc. ent. 41: 9
• minor Nicolesco, 1948; Nota buio. Bucharest 6: 146
• romulanigra Verity, 1950; Le Farfalle diurn. d'Italia 4: 115
• Melitaea didyma ab. lineata Schnaider, 1950; Polskie Pismo. Ent. 19 (3-4): 241
• lobatoi Sabariego, 1976; Revta Lepid. 4 (3): 243

• Didymaeformia didyma; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) didyma; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
• Melitaea (didyma) didyma; [BRU2]: 59, pl. 38, f. 1-12, 25-26
• Melitaea didyma athalia; [LepIndex]; [CoL]
• Melitaea didyma ochracea; [CoL]
• Melitaea didyma; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 429; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [H&R], 116; [BOW]: pl. 3, f. 2-3; [EBW]; [MRS], 586; [Otakar Kudrna]; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43; [FE]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [Mario Maier]
• Melitaea didyma ab. ♀ nigerrima; Schultz, 1908, Ent. Zs. 22 (23): 91
• Melitaea didyma ab. fischeri; Aigner-Abafi, 1906, Ann. hist.-nat. Mus. Hung. 4 (2): 494, pl. 13, f. 4

500x316 (~61Kb) ♀ upperside Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes	500x325 (~44Kb) ♂ upperside Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes	450x432 (~38Kb) underside Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes
597x400 (~49Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	371x550 (~36Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch	597x400 (~52Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch

Larva on Plantago , Veronica , Viola , Scrophularia , Linaria , Dianthus [BRU2]: 59

• M. d. didyma SEU, CEU, W.Siberia, Caucasus, Transcaucasia

  • Melitaea didyma armoricana Oberthür, 1909; Étud. Lépid. Comp. 3: 241
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44
  • Melitaea didyma armoricana f. boulei Oberthür, 1909; Étud. Lépid. Comp. 3: 241, 411, pl. 26, f. 144
  • Melitaea didyma didyma ab. ♂ fulvocincta Bramson, 1910; Horae Soc. ent. Ross. 39: 393
  • Melitaea didyma didyma ab. ♂ demaculata Bramson, 1910; Horae Soc. ent. Ross. 39: 395
  • marginata Ksienschopolski, 1911; Trudy obshch.Izsl. Volyni Zitomir 8: 37, pl. 2, f. 5 (preocc.)
  • latolimbata Sovinsky, 1927; Trav. Sta. biol. Dniepre 2: 381 (repl. marginata Ksienschopolski, 1911)
  • niesiolowskii Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 305
  • cinereoradiata Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 306
  • nigrovenata Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 306
  • uralicola Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 314
  • crimitaea Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 315
  • maledescripta Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 318
  • pontifex Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 321

  • Melitaea didyma armoricana; Eisner, 1942, Zool. Meded. Leiden 24 (4): 118; [LepIndex]; [CoL]
  • Melitaea didyma crimitaea; [LepIndex]; [CoL]
  • Melitaea didyma didyma; [H&R], 116; [BRU2], 59; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44
  • Melitaea didyma maledescripta; [LepIndex]; [CoL]
  • Melitaea didyma niesiolowskii; [LepIndex]; [CoL]
  • Melitaea didyma pontifex; [LepIndex]; [CoL]
  • Melitaea didyma uralicola; [LepIndex]; [CoL]
• M. d. neera Fischer de Waldheim, 1840 SEEU, Volga, S.Ural, SW.Siberia, N.Kazakhstan

  • Melitaea didyma neera Fischer de Waldheim, 1840; Bull. Soc. imp. Nat. Moscou 1840 (1): 81
  • Melitaea didyma var. rossica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.)
  • caucasicola Shlejuzhko, 1940; Folia Zoologica et Hydrobiologica 10 (2): 316, pl. 1, f. 2, pl. 2, f. 12, pl. 6, f. 55
    • = ?Melitaea persea persea; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea uralca Bryk, 1940

  • Melitaea didyma neera; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; [Oleg Kosterin]; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea neera; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43

  820x480 (~92Kb) larva A high steppefied right bank of the Shipunikha Rivulet 3 km NE of the village Evsino, Iskitim district, Novosibirsk Province, West Siberia, Russia. 8th June 1996 (on a leaf of Phlomis tuberosa L.), Photo © Oleg Kosterin

  Larva on Phlomis tuberosa [Oleg Kosterin]

• M. d. orientalis Herrich-Schäffer, 1856

  • Melitaea didyma var. orientalis Herrich-Schäffer, 1856; Syst. Bearb. Schmett. Europ. 6 (48): 4

  • Melitaea didyma orientalis; [LepIndex]; [CoL]
• M. d. occidentalis Staudinger, 1861 Spain

  • Melitaea didyma var. occidentalis Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8; TL: Albarracin, Spain
    • = Melitaea didyma; [FE]
  • Melitaea didyma occidentalis f. dalmatina Staudinger, 1870; Horae Soc. ent. ross. 7 (1870): 60; TL: Greece
  • draconimorta Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 326

  • Melitaea didyma occidentalis; [H&R], 117; [BMAT]: 53, pl. 17, f. 22-36; [LepIndex]; [CoL]
  • Melitaea occidentalis; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 42

  Larva on Linaria spp., Scrophularia sp., Antirrhinum sp., Mispate orontium, Plantago sp., P. amplexicaulis, Melampyrum sp., Verbascum sp., Linaria sagittata [BMAT]

• M. d. alpina Staudinger, 1861

  • Melitaea didyma v. alpina Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8
  • Melitaea didyma var. alpina f. ♀ subalbida Schultz, 1908; Ent. Zs. 22 (23): 91; TL: Wallis
  • Melitaea didyma alpina ab. ♀ latefascia Bramson, 1910; Horae Soc. ent. Ross. 39: 393
  • Melitaea didyma alpina ab. ♀ griseofusca Bramson, 1910; Horae Soc. ent. Ross. 39: 404
  • Melitaea didyma alpina ab. ♀ diluta Bramson, 1910; Horae Soc. ent. Ross. 39: 405

  • Melitaea didyma alpina; [LepIndex]; [CoL]
• M. d. graeca Staudinger, 1870 Greece

  • Melitaea didyma v. graeca Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.)
  • Melitaea didyma var. graeca Staudinger, 1870; Horae Soc. ent. ross. 7 (1870): 60, pl. 1, f. 3

  • Melitaea didyma graeca; [LepIndex]; [CoL]
• M. d. meridionalis Staudinger, 1870 S.France, Alpes Maritimes, Greece, Turkey

  • Melitaea didyma var. meridionalis Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.)
  • Melitaea didyma var. meridionalis Staudinger, 1870; Horae Soc. ent. ross. 7 (1870): 60; TL: Mt. Parnassus, Greece
    • = Melitaea didyma; [FE]
  • Melitaea didyma meridionalis ab. ♀ discolor Bramson, 1910; Horae Soc. ent. Ross. 39: 406
  • patycosana Turati, 1912; Ann. Mus. Napoli 3 (18): 18
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma seilemis Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 9; TL: Alpes Maritimes
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma marsilia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 9; TL: S.France, Riviera
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma marsilia f. androtropia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 10
  • Melitaea didyma marsilia f. ochracea Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 10
  • Melitaea didyma marsilia f. viridescens Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 10
  • Melitaea didyma marsilia f. georgi Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 10; TL: Tessin
  • Melitaea didyma protea Verity, 1917; Bull. Soc. ent. ital. 48: 183
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma subalpina Verity, 1919; Ent. Rec. J. Var. 31: 180
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma protea f. graecaeformis Verity, 1917; Bull. Soc. ent. ital. 48: 183
  • stupenda Nitsche, [1926]; Verh. zool.-bot. Ges. 74/75 (s.b.): 62
  • Melitaea didyma garumna Verity, 1929; Ent. Rec. J. Var. 41 (5): 72; TL: SW.France
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma garumna gen. postgarumna Verity, 1929; Ent. Rec. J. Var. 41 (5): 72; TL: Garzac; St. Laurent d'Arce, Gironde
  • Melitaea didyma subseilemis Verity, 1929; Ent. Rec. J. Var. 41 (5): 73; TL: Mt Aigoual, Cevennes
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma subpatycosana (II) gen. magnaestiva Verity, 1929; Ent. Rec. J. Var. 41 (5): 76
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma microleopardata Verity, 1929; Ent. Rec. J. Var. 41 (5): 76
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45

  • Melitaea didyma androtropia; [LepIndex]; [CoL]
  • Melitaea didyma garumna; [LepIndex]; [CoL]
  • Melitaea didyma georgi; [LepIndex]; [CoL]
  • Melitaea didyma marsilia; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea didyma meridionalis; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [H&R], 116; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea didyma microleopardata; [LepIndex]; [CoL]
  • Melitaea didyma ochracea; [LepIndex]
  • Melitaea didyma patycosana; [LepIndex]; [CoL]
  • Melitaea didyma protea; [LepIndex]; [CoL]
  • Melitaea didyma seilemis; [LepIndex]; [CoL]
  • Melitaea didyma subalpina; [LepIndex]; [CoL]
  • Melitaea didyma subseilemis; [LepIndex]; [CoL]
  • Melitaea didyma viridescens; [LepIndex]; [CoL]
  • Melitaea marsilia; [iNaturalist]
• M. d. crasnensis Hormuzaki, 1894

  • Melitaea didyma var. crasnensis Hormuzaki, 1894; Ent. Nachr. 20 (1): 4; TL: Bukowina

  • Melitaea didyma crasnensis; [LepIndex]; [CoL]
• M. d. kazanskyi Krulikowsky, 1898

  • didyma kazanskyi Krulikowsky, 1898; Mat. pozn. F.F. Ross. Imp. 5: 60
  • ?kazanskyi Krulikowsky, 1901; Mat. k. pozn. F.F. Ross. Imp. 6: 30

  • Melitaea didyma kazanskyi; [LepIndex]; [CoL]
• M. d. nigra Balestre, 1903

  • Melitaea didyma ab. ♂ nigra Balestre, 1903; Bull. Soc. ent. Fr. 1903: 306; TL: nr. Moulinet (Alpes-Maritimes)

  • Melitaea didyma nigra; [LepIndex]; [CoL]
• M. d. mauretanica Oberthür, 1909 N.Africa, S.Spain

  • Melitaea didyma f. mauretanica Oberthür, 1909; Étud. Lépid. Comp. 3: 243; TL: Algeria, Spain
  • Melitaea didyma nisseni Rothschild, 1913; Novit. Zool. 20 (1): 115; TL: Aïn Draham (Tunisia)
  • Melitaea didyma mauretanica (II) gen. occasus Verity, 1919; Ent. Rec. J. Var. 31: 180; TL: [N] Africa
  • Melitaea didyma proteaoccidentis Verity, 1929; Ent. Rec. J. Var. 41 (7-8): 117; TL: Fum Kheney [Foum Kheneg], Mekney [Meknes] (Morocco)
  • Melitaea didyma nisseni gen. tunisa Eisner, 1942; Zool. Meded. Leiden 24 (4): 118; TL: Tunis
  • Melitaea didyma interposita Rothschild, 1913; Novit. Zool. 20 (1): 115; TL: Batna (Algeria)
  • Melitaea didyma africa-felix [= africafelix] Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 326, pl. 1, f. 5; TL: El-Bayadh, Sebdou (Algeria)

  • Melitaea deserticola interposita; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113
  • Melitaea deserticola nisseni; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 114
  • Melitaea didyma africafelix; [LepIndex]; [CoL]
  • Melitaea didyma interposita; [LepIndex]; [CoL]
  • Melitaea didyma mauretanica; [LepIndex]; [CoL]
  • Melitaea didyma nisseni; [LepIndex]; [CoL]
  • Melitaea didyma proteaoccidentis; [LepIndex]; [CoL]
  • Melitaea mauretanica; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43

  600x435 (~43Kb) ♀ upperside Morocco, Djebel Lakraa, Rif, Photo © Michel Tarrier

  600x435 (~28Kb) ♂ upperside Morocco, Itzer, Moyen Atlas, Photo © Jean Delacre

  interposita

  600x435 (~41Kb) underside Morocco, Tizi-n-Test, Haut Atlas, Photo © Michel Tarrier

• M. d. bosphorana Culot, 1909

  • didyma bosphorana Culot, 1909; Bull. Soc. Lép. Genève 1: 166

  • Melitaea didyma bosphorana; [LepIndex]; [CoL]
• M. d. lilliputana Oberthür, 1909 Syria, Armenia, Turkey, Israel

  • Melitaea didyma lilliputana Oberthür, 1909; Étud. Lépid. Comp. 3: 244

  • Melitaea didyma lilliputana; [LepIndex]; [CoL]
  • Melitata liliputana[sic, lilliputana]; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 42
• M. d. castiliana Melcón, 1910

  • Melitaea didyma v. castiliana Melcón, 1910; Bol. Soc. Esp. Hist. nat. 10: 219

  • Melitaea didyma castiliana; [LepIndex]; [CoL]
• M. d. neeraeformis Verity, 1914

  • Melitaea didyma neeraeformis Verity, 1914; Boll. Soc. ent. ital. 45: 212

  • Melitaea didyma neeraeformis; [LepIndex]; [CoL]
• M. d. fasciata Vorbrodt, 1917

  • Melitaea didyma fasciata Vorbrodt, 1917; Mitt. schweiz. ent. Ges. 12 (9-10): 521; TL: Thalgut

  • Melitaea didyma fasciata; [LepIndex]; [CoL]
• M. d. rebrensis Gusic, 1917 Croatia

  • Melitaea didyma rebrensis Gusic, 1917; Godina 29: 216

  • Melitaea didyma rebrensis; [LepIndex]; [CoL]
• M. d. albescens Verity, 1917

  • Melitaea didyma protea f. albescens Verity, 1917; Bull. Soc. ent. ital. 48: 184

  • Melitaea didyma albescens; [LepIndex]; [CoL]
• M. d. druentia Fruhstorfer, 1917 Bosnia

  • Melitaea didyma druentia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 8; TL: Bosnia, Coricna, Maklenpass

  • Melitaea didyma druentia; [LepIndex]; [CoL]
• M. d. lesora Fruhstorfer, 1917

  • Melitaea didyma lesora Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 8; TL: S.Tirol, Klausen, Lana

  • Melitaea didyma lesora; [LepIndex]; [CoL]
• M. d. naina Fruhstorfer, 1917

  • Melitaea didyma naina Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 8; TL: Krain, Laibach
  • didyma postnaina Verity, 1950; Le Farfalle diurn. d'Italia 4: 125

  • Melitaea didyma naina; [LepIndex]; [CoL]
• M. d. oreithyia Fruhstorfer, 1917 Austria

  • Melitaea didyma oreithyia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 7; TL: Austria, Küstenland
  • Melitaea didyma r. oreithyia (II) gen. postoreithyia Verity, 1929; Ent. Rec. J. Var. 41 (6): 91

  • Melitaea didyma oreithyia; [LepIndex]; [CoL]
• M. d. tarlonia Fruhstorfer, 1917

  • Melitaea didyma tarlonia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 8; TL: "Südabhang des Simplon"
  • posttarlonia Verity, 1950; Le Farfalle diurn. d'Italia 4: 128

  • Melitaea didyma tarlonia; [LepIndex]; [CoL]
• M. d. elavar Fruhstorfer, 1917 Ghissar

  • Melitaea didyma elavar Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 11; TL: Dukdan, Sarafschan

  • Melitaea didyma elavar; [BRU2], 59
  • Melitaea didyma elevar; [LepIndex]; [CoL]
• M. d. latefasciata Vorbrodt, 1917

  • Melitaea didyma f. latefasciata Vorbrodt, 1917; Mitt. schweiz. ent. Ges. 12 (9-10): 435

  • Melitaea didyma latefasciata; [LepIndex]; [CoL]
• M. d. subtuscrassipuncta Reverdin, 1919

  • didyma subtuscrassipuncta Reverdin, 1919; Bull. Soc. Lep. Genève 4: pl. 6

  • Melitaea didyma subtuscrassipuncta; [LepIndex]; [CoL]
• M. d. eutitania Fruhstorfer, 1919

  • Melitaea didyma eutitania Fruhstorfer, 1919; Archiv Naturg. 83 A (6): 13; TL: Liguria, Pegli

  • Melitaea didyma eutitania; [LepIndex]; [CoL]
• M. d. nigrorubida Verity, 1919

  • Melitaea didyma nigrorubida Verity, 1919; Ent. Rec. J. Var. 31: 180; TL: Saxony

  • Melitaea didyma nigrorubida; [LepIndex]; [CoL]
• M. d. subrubida Verity, 1919

  • Melitaea didyma subrubida Verity, 1919; Ent. Rec. J. Var. 31: 180

  • Melitaea didyma subrubida; [LepIndex]; [CoL]
• M. d. apenninigena Verity, 1919 Apennines

  • Melitaea didyma apenninigena Verity, 1919; Ent. Rec. J. Var. 31: 181

  • Melitaea didyma apenninigena; [LepIndex]; [CoL]
• M. d. palustris Verity, 1919 Italy

  • Melitaea didyma palustris Verity, 1919; Ent. Rec. J. Var. 31: 181

  • Melitaea didyma palustris; [LepIndex]; [CoL]
• M. d. roccii Turati, 1921

  • Melitaea didyma roccii Turati, 1921; Naturalista sicil. 23 (7-12): 225

  • Melitaea didyma roccii; [LepIndex]; [CoL]
• M. d. leopardata Verity & Querci, 1924 Italy

  • Melitaea didyma f. leopardata Verity & Querci, 1924; Ent. Rec. (suppl.) 36: 37

  • Melitaea didyma leopardata; [LepIndex]; [CoL]
• M. d. subpatycosana Verity & Querci, 1924 Italy

  • Melitaea didyma subpatycosana Verity & Querci, 1924; Ent. Rec. (suppl.) 36: 38

  • Melitaea didyma subpatycosana; [LepIndex]; [CoL]
• M. d. alaeida Stauder, 1925 Tyrol

  • Melitaea didyma f. alaeida Stauder, 1925; Ent. Anz. 5 (6): 43

  • Melitaea didyma alaeida; [LepIndex]; [CoL]
• M. d. supercaldaria de Sagarra, 1926 Spain

  • didyma supercaldaria de Sagarra, 1926; Butll. Inst. catal. Hist. nat. (2) 6 (6-7): 130

  • Melitaea didyma supercaldaria; [LepIndex]; [CoL]
• M. d. inversa Skala, 1928

  • Melitaea didyma ab. inversa Skala, 1928; Z. Öst. EntVer 13: 85

  • Melitaea didyma inversa; [LepIndex]; [CoL]
• M. d. mariannae Skala, 1928

  • Melitaea didyma mariannae Skala, 1928; Z. Öst. EntVer 13: 85; TL: Liegnitz

  • Melitaea didyma mariannae; [LepIndex]; [CoL]
• M. d. calcea Skala, 1928

  • Melitaea didyma var. calcea Skala, 1928; Z. Öst. EntVer 13: 87

  • Melitaea didyma calcea; [LepIndex]; [CoL]
• M. d. parvalpina Verity, 1928

  • Melitaea didyma r. parvalpina Verity, 1928; Ent. Rec. 40: 163

  • Melitaea didyma parvalpina; [LepIndex]; [CoL]
• M. d. cataprotea Verity, 1929 Spain

  • Melitaea didyma gen. cataprotea Verity, 1929; Ent. Rec. J. Var. 41 (7-8): 117
  • Melitaea didyma gen. cataminuta Verity, 1929; Ent. Rec. J. Var. 41 (7-8): 118; TL: Vallvidrena; La Garriga
  • Melitaea didyma ab. hiberava Verity, 1929; Ent. Rec. J. Var. 41 (9): 130; TL: Orihuela, Aragon
  • Melitaea didyma gen. cataoccasus Verity, 1929; Ent. Rec. J. Var. 41 (7-8): 118

  • Melitaea didyma catoprotea; [LepIndex]; [CoL]
• M. d. rectealpina Verity, 1929

  • Melitaea didyma rectealpina Verity, 1929; Ent. Rec. J. Var. 41 (4): 66; TL: Bormio

  • Melitaea didyma rectealpina; [LepIndex]; [CoL]
• M. d. superalpina Verity, 1929

  • Melitaea didyma superalpina Verity, 1929; Ent. Rec. J. Var. 41 (4): 66; TL: Val di Fraele
• M. d. subtarlonia Verity, 1929 Italy

  • Melitaea didyma subtarlonia Verity, 1929; Ent. Rec. J. Var. 41 (5): 73; TL: Pian del Sole, mtns above Intra, lake Maggiore
  • Melitaea didyma gen. praemarsilia Verity, 1929; Ent. Rec. J. Var. 41 (5): 74
  • Meliaea didyma ab. atrarubra Verity, 1929; Ent. Rec. J. Var. 41 (5): 75
  • georginigrior Verity, 1950; Le Farfalle diurn. d'Italia 4: 124

  • Melitaea didyma subtarlonia; [LepIndex]; [CoL]
• M. d. pyrenealpina Verity, 1929 Pyrenees

  • Melitaea didyma pyrenealpina Verity, 1929; Ent. Rec. J. Var. 41 (5): 74; TL: Gèdre, 1100m, Hautes Pyrénées

  • Melitaea didyma pyrenealpina; [LepIndex]; [CoL]
• M. d. atralpina Verity, 1929

  • Melitaea didyma atralpina Verity, 1929; Ent. Rec. J. Var. 41 (6): 89

  • Melitaea didyma atralpina; [LepIndex]; [CoL]
• M. d. ravalpina Verity, 1929 Tyrol

  • Melitaea didyma ravalpina Verity, 1929; Ent. Rec. J. Var. 41 (6): 90

  • Melitaea didyma ravalpina; [LepIndex]; [CoL]
• M. d. turkestanica Sheljuzhko, 1929 Kazakhstan, Kyrgyzstan, Uzbekistan, Tajikistan

  • Melitaea didyma turkestanica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 354 (repl. Melitaea turanica Staudinger, 1886)
  • Melitaea didyma var. turanica Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 231 (preocc.)
  • Melitaea didyma turkestanica ab. pseudoala Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 355; TL: Kauntshi, Tashkent
    • = ; [Wahlberg]
  • Melitaea didyma turkestanica ab. alboocellata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 355

  • Didymaeformia pseudoala; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
  • Melitaea didyma turkestanica; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [BRU2], 59; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; Yakovlev, 2012, Nota lepid. 35 (1): 88; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea didyma var. turanica; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 429
  • Melitaea pseudoala; [LepIndex] ; [CoL]
  • Melitaea turkestanica; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43

  Misidentified/misapplied:

  • Melitaea pseudoala; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 245, pl. 4, f. 12, pl. 16, f. 4
    • = Melitaea ninae; Kolesnichenko, 1999, Atalanta 30 (1/4): 101
  • Melitaea pseudoala; Higgins, 1955, Trans. R. ent. Soc. Lond. 106 (1): 116
    • = Melitaea ninae; Kolesnichenko, 1999, Atalanta 30 (1/4): 101
  • Melitaea pseudoala; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166
    • = Melitaea ninae; Kolesnichenko, 1999, Atalanta 30 (1/4): 101
  • Melitaea pseudoala; Korshunov, 1972, Ent. Obozr. 51 (2): 355
    • = Melitaea ninae; Kolesnichenko, 1999, Atalanta 30 (1/4): 101
  • Melitaea pseudoala; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
    • = Melitaea ninae; Kolesnichenko, 1999, Atalanta 30 (1/4): 101
• M. d. libanotica Belter, 1934 Lebanon

  • Melitaea didyma libanotica Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 106, f. 1-2; TL: Bscharre, N.Lebanon, Syria, 1300m; Zahatta
  • ibra Higgins, 1964; Entomologist 97: 196
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44
  • schwingenschussi Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 323
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44

  • Melitaea didyma ibra; [LepIndex]; [CoL]
  • Melitaea didyma libanotica; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea didyma schwingenschussi; [LepIndex]; [CoL]
• M. d. variabilis Belter, 1934 Sicily

  • Melitaea didyma variabilis Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 108; TL: Karaabey
  • Melitaea didyma sicula Stauder, 1928; Lep. Rdsch. 2 (10): 107 (preocc.)
  • Melitaea didyma variablilis gen. aest. aestiva Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 109

  • Melitaea didyma variabilis; [LepIndex]; [CoL]
• M. d. tschecha Bryk, 1940

  • didyma tschecha Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 301

  • Melitaea didyma tschecha; [LepIndex]; [CoL]
• M. d. austria Bryk, 1940

  • didyma austria Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 302

  • Melitaea didyma austria; [LepIndex]; [CoL]
• M. d. magnalpina Bryk, 1940

  • didyma magnalpina Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 304

  • Melitaea didyma magnalpina; [LepIndex]; [CoL]
• M. d. gravosana Bryk, 1940

  • didyma gravosana Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 307

  • Melitaea didyma gravosana; [LepIndex]; [CoL]
• M. d. embriki Bryk, 1940

  • didyma embriki Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 311

  • Melitaea didyma embriki; [LepIndex]; [CoL]
• M. d. irridens Bryk, 1940

  • didyma irridens Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 312

  • Melitaea didyma irridens; [LepIndex]; [CoL]
• M. d. kirgisica Bryk, 1940 Tarbagatai, Saur, Tian-Shan

  • didyma kirgisica Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 315
  • geminella Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 332

  • Melitaea didyma geminella; [LepIndex]; [CoL]
  • Melitaea didyma kirgisica; [BRU2], 59
• M. d. walteri Bryk, 1940 Mongolia

  • didyma walteri Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 343

  • Melitaea didyma walteri; [LepIndex]; [CoL]
• M. d. ambra Higgins, 1941

  • didyma ambra Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 208 (repl. Melitaea didyma dschungarica Oberthür, 1909)
  • Melitaea didyma dschungarica Oberthür, 1909; Étud. Lépid. Comp. 3: 243 (preocc.)
  • perplexa Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 239
  • Melitaea didyma manoni Hanus, 1996; Atalanta 27 (1/2): 207; TL: Kirghizia, Kadzhi Say, 1610m
    • = Melitaea didyma turkestanica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38

  • Melitaea didyma ambra; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • ambra; [BRU2], 59 (?)
• M. d. cinerea Higgins, 1941 Sicily, Asia Minor, N.Syria

  • didyma cinerea Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 209

  • Melitaea didyma cinerea; [LepIndex]; [CoL]
• M. d. boschmai Eisner, 1942 Macedia

  • Melitaea didyma boschmai Eisner, 1942; Zool. Meded. Leiden 24 (4): 117; TL: Monastir, Prilep, Olympus, Saloniki (Macedonien)

  • Melitaea didyma boschmai; [LepIndex]; [CoL]
• M. d. orinomus Rütimeyer, 1942 Pyrenees

  • didyma orinomus Rütimeyer, 1942; Mitt. schweiz. ent. Ges. 18 (9-10): 435

  • Melitaea didyma orinomus; [LepIndex]; [CoL]
• M. d. ochra Verity, 1950 Italy

  • didyma ochra Verity, 1950; Le Farfalle diurn. d'Italia 4: 104

  • Melitaea didyma ochra; [LepIndex]; [CoL]
• M. d. celladdita Verity, 1950 Italy

  • didyma celladdita Verity, 1950; Le Farfalle diurn. d'Italia 4: 106

  • Melitaea didyma celladdita; [LepIndex]; [CoL]
• M. d. cellapartita Verity, 1950 Italy

  • didyma cellapartita Verity, 1950; Le Farfalle diurn. d'Italia 4: 106

  • Melitaea didyma cellapartita; [LepIndex]; [CoL]
• M. d. medioduplicata Verity, 1950 Italy

  • didyma medioduplicata Verity, 1950; Le Farfalle diurn. d'Italia 4: 106

  • Melitaea didyma medioduplicata; [LepIndex]; [CoL]
• M. d. postcelladdita Verity, 1950 Italy

  • didyma postcelladdita Verity, 1950; Le Farfalle diurn. d'Italia 4: 106

  • Melitaea didyma postcelladdita; [LepIndex]; [CoL]
• M. d. deletasicca Verity, 1950 Italy

  • didyma deletasicca Verity, 1950; Le Farfalle diurn. d'Italia 4: 107

  • Melitaea didyma deletasicca; [LepIndex]; [CoL]
• M. d. contracta Verity, 1950 Italy

  • didyma contracta Verity, 1950; Le Farfalle diurn. d'Italia 4: 108

  • Melitaea didyma contracta; [LepIndex]; [CoL]
• M. d. lucida Verity, 1950 Italy

  • didyma lucida Verity, 1950; Le Farfalle diurn. d'Italia 4: 108

  • Melitaea didyma lucida; [LepIndex]; [CoL]
• M. d. porrecta Verity, 1950 Italy

  • didyma porrecta Verity, 1950; Le Farfalle diurn. d'Italia 4: 108

  • Melitaea didyma porrecta; [LepIndex]; [CoL]
• M. d. folvonigrolimbata Verity, 1950 Italy

  • didyma folvonigrolimbata Verity, 1950; Le Farfalle diurn. d'Italia 4: 110

  • Melitaea didyma folvonigrolimbata; [LepIndex]; [CoL]
• M. d. infracrasselunata Verity, 1950 Italy

  • didyma infracrasselunata Verity, 1950; Le Farfalle diurn. d'Italia 4: 110

  • Melitaea didyma infracrasselunata; [LepIndex]; [CoL]
• M. d. inframedioconfluens Verity, 1950 Italy

  • didyma inframedioconfluens Verity, 1950; Le Farfalle diurn. d'Italia 4: 110

  • Melitaea didyma inframedioconfluens; [LepIndex]; [CoL]
• M. d. posticemediocrasse Verity, 1950 Italy

  • didyma posticemediocrasse Verity, 1950; Le Farfalle diurn. d'Italia 4: 110

  • Melitaea didyma posticemediocrasse; [LepIndex]; [CoL]
• M. d. siccalta Verity, 1950

  • didyma siccalta Verity, 1950; Le Farfalle diurn. d'Italia 4: 116

  • Melitaea didyma siccalta; [LepIndex]; [CoL]
• M. d. lesoralpina Verity, 1950 Italy

  • didyma lesoralpina Verity, 1950; Le Farfalle diurn. d'Italia 4: 128

  • Melitaea didyma lesoralpina; [LepIndex]; [CoL]
• M. d. pseudolatonigena Yakovlev, 2002 Tuva

  • Melitaea didyma pseudolatonigena Yakovlev, 2002; Helios 3: (102-106)
    • = Melitaea didyma altaica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38

Melitaea interrupta Kolenati, 1846 Caucasus, Turkey, Iran. See [maps]

• Melitaea interrupta Kolenati, 1846; Meletamata Entomologica 5: 86; TL: "Caucaso et Transcaucasia"

• Melitaea (Didymaeformia) interrupta; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
• Melitaea (didyma) interrupta; [BRU2]: 60, pl. 38, f. 13-18
• Melitaea interrupta; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43; [iNaturalist] ; [LEPIFORUM]

• M. i. interrupta NE.Turkey?, Caucasus, Transcaucasus

  • Melitaea didyma transcaucasica Turati, 1921; Naturalista sicil. 23 (7-12): 225
    • = Melitaea interrupta; [Wahlberg]
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea trivia caucasi Verity, 1922; Ent. Rec. J. Var. 34: 13
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma venosata Niesiolowski, 1937; Ann. Mus. zool. polon. 11 (28): 469; TL: Caucasus
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • distinctissima Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 345
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • musatscherii Alberti, 1969
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • itkolensis Alberti, 1969
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45

  • Didymaeformia transcaucasica; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
  • Melitaea didyma caucasi; Verity, 1929, Ent. Rec. J. Var. 41 (3): 43, (7-8):112
  • Melitaea didyma distinctissima; [LepIndex]; [CoL]
  • Melitaea didyma transcaucasica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38
  • Melitaea interrupta interrupta; [BRU2], 60; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea transcaucasica; [LepIndex] ; [CoL] ; [iNaturalist]
  • Melitaea trivia caucasi; [BRU2], 62; [iNaturalist]
• M. i. kendevana Gross & Ebert, 1975 Kopet-Dagh

  • Melitaea transcaucasica maculata Gross & Ebert, 1975; J. ent. Soc. Iran, suppl. 1: 40 (preocc. Melitaea astreoidea maculata Staudinger, 1901)
  • elbursicola Gross & Ebert, 1975
  • Melitaea transcaucasica grossi Koçak, 1980; Nota lepid. 2 (4): 141 (repl. Melitaea transcaucasica maculata Gross & Ebert, 1975)

  • Melitaea interrupta kendevana; [BRU2], 60; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea transcaucasica grossi; [LepIndex]; [CoL]
  • Melitaea transcaucasica maculata; [LepIndex]; [CoL]

Melitaea latonigena Eversmann, 1847 ..., NE.Kazakhstan, Mongolia, NW.China. See [maps]

• Melitaea latonigena Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (3): 66, pl. 1, f. 1-2; TL: [Irkutsk Region, Russia]
• mongoligena Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 342

• Melitaea (Didymaeformia) latonigena; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea (didyma) latonigena; [BRU2]: 60, pl. 37, f. 31-36
• Melitaea didyma latonigena; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [LepIndex]; [CoL]
• Melitaea didyma latonigera[sic]; [MRS], 586
• Melitaea didyma mongoligena; [LepIndex]; [CoL]
• Melitaea latonigena; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43; [iNaturalist] ; [SZMN]

• M. l. latonigena Tuva - Baikal, Sayan

  • latonigena atrata Higgins, 1935; Entomologist 68: 112
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

  • Melitaea latonigena latonigena; [BRU2], 60; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

  atrata

  1140x782 (~172Kb) ♂ ♀ Russia, Siberia, Tyva Republic, Todzha District, the Azas Nature Reserve, a southern steppen slope of a mountain on the northern bank of Lake Azas at the Ilgi-Chul brook mouth, 52°44' N 96°36' E, alt. 1000 m, 23rd July 2000, Photo © Oleg Kosterin

• M. l. altaica Grum-Grshimailo, 1893 Altai

  • Melitaea didyma var. altaica Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 (1-2): 127; TL: SW.Altai, Lake Markakol

  • Melitaea didyma altaica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 38; [LepIndex]; [CoL]
  • Melitaea latonigena altaica; [BRU2], 60; [Oleg Kosterin]; Yakovlev, 2012, Nota lepid. 35 (1): 88

  461x700 (~58Kb) ♂ ♀ underside A subalpine short forb meadow on an opening in an open larch forest on the southern principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets , 2100 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16th July 1998, Photo © Oleg Kosterin	400x533 (~49Kb) ♂ underside An alpine meadow, at 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 14th July, 1988, Photo © Oleg Kosterin	429x668 (~28Kb) underside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin
  768x700 (~76Kb) underside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin	572x407 (~28Kb) ♀ upperside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998, Photo © Oleg Kosterin	503x432 (~21Kb) ♂ (aberrative) underside A steppefied southern slope of small mountains on the Dzhazator River right bank in front of the Akbul Rivulet mouth, between the Yuzhno-Chuiskii mountain range and the Ukok Plateau, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 19th July 1998, Photo © Oleg Kosterin
  1099x852 (~410Kb) ♂ upperside The lower Oroktoi brook valley, a right tributary of the Akkem River, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 7th July 2007 (on fresh horse dung), Photo © Oleg Kosterin

• M. l. polaris Grum-Grshimailo, 1899 Yakutia

  • ?Melitaea didyma var. polaris Grum-Grshimailo, 1899; Ann. Mus. Zool. Imp. Acad. Sci. St.-Petersb. 4 (4): 459

  • Melitaea didyma polaris; [LepIndex]; [CoL]
  • Melitaea latonigena polaris; [BRU2], 60; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. l. ukoka Korshunov, 1998

  • Melitaea latonigena ukoka; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39; [SZMN]

Melitaea mixta Evans, 1912 Tajikistan, Pakistan, Afghanistan. See [maps]

• Melitaea didyma mixta Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 583, 558 [?]; TL: "Chit.[ral]"

• Didymaeformia mixta; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (didyma) mixta; [BRU2]: 61, pl. 41, f. 1-3
• Melitaea didyma mixta; ; [LepIndex]; [CoL]
• Melitaea mixta; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43

• M. m. mixta NW.India

  • Melitaea mixta mixta; [BRU2], 61

Melitaea nadezhdae Sheljuzhko, 1912 -. See [maps]

• Melitaea didyma nadezhdae Sheljuzhko, 1912; Dt. ent. Z. Iris 26 (3): 137; TL: Pamirskij-Post

• Melitaea (Didymaeformia) nadezhdae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea didyma nadezhdae; [LepIndex]; [CoL]
• Melitaea mixta nadezhdae; [BRU2], 61

• M. n. nadezhdae E.Pamirs

  • Melitaea nadezhdae nadezhdae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. n. problematica Sheljuzhko, 1929 W.Pamirs

  • Melitaea didyma f. problematica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 357, pl. 27, f. 11-12

  • Melitaea mixta problematica; [BRU2], 61
  • Melitaea nadezhae problematica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

Melitaea persea Kollar, [1850] Iran, Asia Minor - Afghanistan, W.Tien-Shan. See [maps]

• Melitaea persea Kollar, [1850]; Denkschr. Akad. Wiss. Wien. 1: 52; TL: "Farsistan" [Shiraz, Iran]
• Melitaea didyma var. romana Calberla, 1887; Corr.-Bl. Ent. Ver. Iris 1 (1) (4): 132
• Melitaea casta var. afghana Heydemann, [1955]; Zs. Wien. Ent. Ges. 39 (12): 416, pl. 35, f. 1-3
  • = ; [BRU2], 61

• Didymaeformia afghana; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Didymaeformia persea; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) persea; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea (didyma) persea; [BRU2]: 61, pl. 44, f. 1-9
• Melitaea casta afghana; [LepIndex]; [CoL]
• Melitaea didyma persea; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113
• Melitaea didyma var. persea; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 432
• Melitaea persea; Weiss & Major, 2000, Atalanta 31 (1/2): 123 (note), pl. VII, f. 9-16; [AfrL]; Carbonell, 2007, Bull. Soc. ent. Fr. 112 (3): 334 (note); Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

• M. p. persea Iraq, Turkey, Jordan

  • Melitaea didyma araratica Verity, 1929; Ent. Rec. J. Var. 41 (7-8): 113
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma magnacasta Verity, 1929; Ent. Rec. J. Var. 41 (9): 128
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea tauricus Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 109; TL: Marasch, N.Syria
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea tauricus gen. aest. microtauricus Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 110
    • = Melitaea persea; [BRU2], 61
  • hafiz Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 225
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • darius Gross & Ebert, 1975
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea hafiz pfeifferi Gross & Ebert, 1975; J. ent. Soc. Iran, suppl. 1: 37 (preocc. Melitaea didyma pfeifferi Bryk, 1940)
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • wernickei Gross & Ebert, 1975
    • = Melitaea persea; [BRU2], 61
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • safawides Gross & Ebert, 1975
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea hafiz eberti Koçak, 1980; Nota lepid. 2 (4): 141 (repl. Meliaea hafiz pfeifferi Gross & Ebert, 1975)

  • Melitaea (Cinclidia) hafiz; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42
  • Melitaea eberti; [iNaturalist] ; [LEPIFORUM]
  • Melitaea hafiz eberti; [LepIndex]; [CoL]
  • Melitaea hafiz; [LepIndex] ; [CoL]
  • Melitaea persea araratica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea persea persea; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea persea tauricus; [LepIndex]; [CoL] ; [iNaturalist]
• M. p. caucasica Staudinger, 1861 Caucasus

  • Melitaea didyma var. caucasica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8
    • = Melitaea persea; [BRU2], 61
    • = Melitaea persea persea; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea didyma var. kaschtschenkoi Christoph, 1889; Ent. Nachr. 15 (5): 69
    • = Melitaea persea; [BRU2], 61

  • Melitaea didyma caucasica; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113
  • Melitaea didyma var. caucasica; Staudinger, 1871, Cat. Lepid. eur. Faunegeb. (Edn. 2): 18
  • Melitaea persea caucasica; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea trivia caucasica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• M. p. dodgsoni Grose-Smith, 1887 Afghanistan

  • Melitaea dodgsoni Grose-Smith, 1887; Ann. Mag. nat. Hist. (5) 20: 130; TL: Gwashki
    • = Melitaea persea; [BRU2], 61

  • Melitaea persea dodgsoni; [CoL]
• M. p. paphlagonia Fruhstorfer, 1917 Transcaspia

  • Melitaea didyma paphlagonia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 10; TL: Arwas, Transcaspia
    • = Melitaea persea; [BRU2], 61

  • Melitaea persea paphlagonia; [LepIndex]; [CoL] ; [iNaturalist]
• M. p. montium Belter, 1934 Syria, Lebanon, Israel

  • Melitaea montium Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 107; TL: Bscharre, N.Lebanon
    • = Melitaea persea; [BRU2], 61

  • Melitaea montium; [LepIndex] ; [CoL]
  • Melitaea persea montium; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
• M. p. sargon Hemming, 1932 Iraq

  • persea sargon Hemming, 1932; Trans. ent. Soc. Lond. 80: 272
    • = Melitaea persea; [BRU2], 61
    • = Melitaea persea persea; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45

  • Melitaea persea sargon; [AfrL]
• M. p. sangdana Eckweiler, 1981 S.Iran

  • Melitaea persea sangdana Eckweiler, 1981; Nachr. ent. Ver. Apollo NF 2 (1): 47; TL: S.Iran, Kerman, 2800-3000m

Melitaea deserticola Oberthür, 1909 N.Africa, Tripolitania, Egypt, Lebanon, Transjordan. See [maps] Desert Fritillary

• Melitaea deserticola Oberthür, 1909; Étud. Lépid. Comp. 3: 243
• Melitaea ab. deserticola Oberthür, 1876; Étud. d'Ent. 1: 25, pl. 3, f. 1; TL: Biskra (Algeria)

• Didymaeformia deserticola; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea deserticola; [H&R], 117; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [AfrL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]

600x435 (~35Kb) upperside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre	600x435 (~68Kb) ♀ upperside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre	600x435 (~59Kb) underside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre
600x435 (~53Kb) upperside Morocco, Aït-Abdallah, Anti-Atlas , Photo © Michel Tarrier

• M. d. deserticola Tunisia, Algeria, Morocco

  • Melitaea didyma harterti Rothschild, 1913; Novit. Zool. 20 (1): 115; TL: El Hadadra, El Golea, Ghardaia (Algeria)
  • Melitaea didyma harterti ab. pseudoacraeina Rothschild, 1917; Novit. Zool. 24 (1): 102; TL: El Hadadra (Algeria)
  • Melitaea didyma harterti ab. nigrofasciata Rothschild, 1917; Novit. Zool. 24 (1): 102; TL: El Hadadra (Algeria)

  • Melitaea deserticola deserticola; [BMAT]: 54, pl. 17, f. 37-38, pl. 18, f. 1-7
  • Melitaea deserticola harterti; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 114

  Larva on Linaria aegyptiaca, Plantago media, Anarrhinum fruticosum, (in Lebanon) Anarrhinum spp. [BMAT]

• M. d. macromaculata Belter, 1934 Lebanon

  • Melitaea deserticola macromaculata Belter, 1934; Arb. Morph. Taxon. Ent. Berl. 1 (2): 106; TL: N. Ibrahim, N.Lebanon
  • belteri Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 324
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44
  • warneckei Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 324
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44

  • Melitaea deserticola macromaculata; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 44; [AfrL]; [LepIndex]; [CoL] ; [iNaturalist]
  • Melitaea didyma warneckei; [LepIndex]; [CoL]
• M. d. scotti Higgins, 1941 Yemen

  • Melitaea abyssinica scotti Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 236

  • Melitaea abyssinica scotti; [LepIndex]; [CoL]
  • Melitaea deserticola scotti; [AfrL]; [iNaturalist]

Melitaea didymina Staudinger, 1895 Mongolia. See [maps]

• Melitaea didymina Staudinger, 1895; Dt. Ent. Z. Iris 8 (2): 347

• Didymaeformia didymina; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) didymina; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea didymina; Yakovlev, 2012, Nota lepid. 35 (1): 88; [LepIndex] ; [CoL] ; [iNaturalist]

• M. d. higginsi Sakai, 1978 Afghanistan

  • Melitaea didymina higginsi Sakai, 1978; Atalanta 9 (1): 116; TL: Bala-Ouran, 3500m, Anjuman valley, NE. Hindukush Mts, Afghanistan

  • Melitaea didymina higginsi; [LepIndex]; [CoL]
  • Melitaea higginsi; [iNaturalist]

Melitaea didymoides Eversmann, 1847 S.Siberia, Mongolia, N.China. See [maps]

• Melitaea didymoides Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (3): 67, pl. 1, f. 3-4; TL: [Kyakhta, Buryatia]
  • = Melitaea ala; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113
• Melitaea didyma mandschurica Seitz, 1909; Gross-Schmett. Erde 1: 219, pl. 66 e (preocc.)
  • = Melitaea ala; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113
• seitzi Matsumura, 1929; Ill. comm. Ins. Japan 1: 15 (repl. Melitaea didyma mandschurica Seitz, 1909)
• mandschukoana Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 336

• Didymaeformia didymoides; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) didymoides; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea didyma mandschukoana; [LepIndex]; [CoL]
• Melitaea didymoides; [MRS], 586; [BRU2]: 62, pl. 39, f. 19-24; Yakovlev, 2012, Nota lepid. 35 (1): 88; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43; [LepIndex] ; [CoL] ; [iNaturalist]

592x710 (~76Kb) ♂ underside A steppefied meadow at Lake Butyvken, surrounded by a magnificent pine forest within a steppe zone called Tsasucheiskii Bor, 10 km SW of the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 4th July 1996, Photo © Oleg Kosterin

800x800 (~153Kb) ♀ upperside a meadow on the Onon River right bank floodland 10 km upstream of the village Nizhnii Tsasuchei, SE Zabaikalye, 18th July 1997 (on Dianthus versicolor), Photo © Oleg Kosterin

• M. d. didymoides S.Tuva?, Transbaikalia, Amur

  • Melitaea didymoides didymoides; [BRU2], 62
• M. d. latonia Grum-Grshimailo, 1891 C.China

  • Melitaea didyma var. latonia Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 455

  • Melitaea didymoides latonia; [MRS], 586; [BRU2], 62
• M. d. pekinensis Seitz, 1909 N.China

  • Melitaea didyma pekinensis Seitz, 1909; Gross-Schmett. Erde 1: 219, pl. 66 e

  • Melitaea didymoides pekinensis; [MRS], 586; [BRU2], 63
  • Melitaea pekinensis; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [LepIndex] ; [CoL]
• M. d. eupatides Fruhstorfer, 1917 Gansu

  • Melitaea didyma eupatides Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 11; TL: Kansu
    • = Melitaea ala; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113
  • didyma kansicola Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 338

  • Melitaea didyma kansicola; [LepIndex]; [CoL]
  • Melitaea didymoides eupatides; [BRU2], 63
• M. d. yugakuana Matsumura, 1927 S.Ussuri

  • Melitaea didyma yugakuana Matsumura, 1927; Insecta Matsumurana 1 (4): 164; TL: S.Manchuria, Prov. Yugakujo

  • Melitaea didymoides yugakuana; [BRU2], 62
  • Melitaea yugakuana; [iNaturalist]
• M. d. hummeli Bryk, 1940 S.Mongolia

  • didymoides hummeli Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 338

  • Melitaea didyma hummeli; [LepIndex]; [CoL]
  • Melitaea didymoides hummeli; [BRU2], 63

Melitaea ala Staudinger, 1881 Tian-Shan, Dzhungarsky Ala-Tau Mts., S.Altai. See [maps]

• Melitaea didyma var. ala Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 288; TL: Lepsa, Ala-Tau

• Didymaeformia ala; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) ala; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea ala; Belter, 1934, Arb. Morph. Taxon. Ent. Berl. 1 (2): 113; [BRU2]: 63, pl. 39, f. 10-15; Kolesnichenko, 1999, Atalanta 30 (1/4): 91; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

Misidentified/misapplied:

• Melitaea didyma var. ala; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 430
  • = Melitaea kotshubeji kotshubeji; Kolesnichenko, 1999, Atalanta 30 (1/4): 96

Larva on Ligularia [BRU2]: 63

• M. a. ala S.Altai, Saur Mts., Dzhungarsky Ala-Tau, Burchansarytau Mts., Tyshkantau Mts., Kaiandy-Tau, Horgos R., Alty-Emel Mts.

  • Melitaea ala ab. immodulata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 366; TL: Dzharkent
    • = ; Kolesnichenko, 1999, Atalanta 30 (1/4): 92
  • Melitaea ala allah Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 346, pl. 3, f. 23; TL: "Siberia"
    • = ; Kolesnichenko, 1999, Atalanta 30 (1/4): 92

  • Melitaea ala ala; Sheljuzhko, 1929, Mitt. Münch. Ent. Ges. 19: 368; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 346; Kolesnichenko, 1999, Atalanta 30 (1/4): 91, pl. 6, f. 1-4; [BRU2], 63; Kolesnichenko, 1999, Atalanta 30 (1/4): 91; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea ala allah; [LepIndex]; [CoL]
  • Melitaea ala; Sushkin, 1913, Z. wiss. Insektenbiologie 9 (6/7): 171, f. 5-6; Wagner, 1913, Ent. Mitt. 2 (3): 91-93; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 242-244, pl. 2, f. 4, 10; Lukhtanov & Lukhtanov, 1994, Herbipoliana 3: 184, pl. 35, f. 5-6

  Misidentified/misapplied:

  • Melitaea ala; Lukhtanov & Lukhtanov, 1994, Herbipoliana 3: pl. 35, f. 5-6
    • = Melitaea ala zaisana; Lukhtanov, 1999, Atalanta 30 (1/4): 144
• M. a. bicolor Seitz, 1907 Kungey Ala-Tau, Kirgizsky Mts., Suusamyr Valley, Suusamyr Mts., Talassky Ala-Tau, Terskey Ala-Tau, Baidullu Mts., Naryn-Too Mts.

  • Melitaea ala ab. bicolor Seitz, 1907; Gross-Schmett. Erde 1: 219, pl. 66, f. 7-8; TL: Karagatai Mts.
    • = Melitaea ala ala; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea ala ab. latemarginata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 366, pl. 28, f. 7-8; TL: Naryn
    • = ; Kolesnichenko, 1999, Atalanta 30 (1/4): 93, pl. 6, f. 5-7

  • Melitaea ala bicolor; Sushkin, 1913, Z. wiss. Insektenbiologie 9 (6/7): 172; Sheljuzhko, 1929, Mitt. Münch. Ent. Ges. 19: 368; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 347-348; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 244, pl. 6, f. 6, 12; Kolesnichenko, 1999, Atalanta 30 (1/4): 93, pl. 6, f. 5-7; [BRU2], 63; [LepIndex]; [CoL]

  Misidentified/misapplied:

  • Melitaea ala bicolor; Shchetkin, 1981, Ent. Obozr. Tajikistana: 163-164
    • = Melitaea kotshubeji kotshubeji; Kolesnichenko, 1999, Atalanta 30 (1/4): 96

  Larva on Ligularia thompsonii Shchetkin, 1981, Ent. Obozr. Tajikistana

• ?M. a. sheljuzhkoi Bryk, 1940 E.Tian-Shan

  • ala sheljuzhkoi Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 348

  • Melitaea ala sheljuzhkoi; [BRU2], 63 (?); [LepIndex]; [CoL]
• ?M. a. determinata Bryk, 1940 E.Tian-Shan

  • ala determinata Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 349

  • Melitaea ala determinata; [BRU2], 63 (?); [LepIndex]; [CoL]
• ?M. a. strandi Bryk, 1940 E.Tian-Shan

  • ala strandi Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 350

  • Melitaea ala strandi; [BRU2], 63 (?); [LepIndex]; [CoL]
• M. a. elleclaudia Bryk, 1940 Kazakhstan

  • ala elleclaudia Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 351
• M. a. zaisana Lukhtanov, 1999 S.Altai

  • Melitaea ala zaisana Lukhtanov, 1999; Atalanta 30 (1/4): 143; TL: Kazakstan, Ust-Kamenogorsk, S.Altai, between Kurtschum and Kalguty, 600m

  • Melitaea ala zaisana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. a. irtyschica Lukhtanov, 1999 W.Altai

  • Melitaea ala irtyschica Lukhtanov, 1999; Atalanta 30 (1/4): 144; TL: Kazakstan, Ust-Kamenogorsk, Ulbinski, Oktjabrsk, 420m

Melitaea kotshubeji Sheljuzhko, 1929 Peter the Great Mts., N.Hazreti-Sho Mts., W.Zaalaisky Mts., S slopes (Alaisky Mts.), SW slopes (Fergansky Mts.), SE slopes (Cathkalsky Mts.). See [maps]

• Melitaea ala kotshubeji Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 364, pl. 27, f. 1-4; TL: Touptshek

• Melitaea (Didymaeformia) kotshubeji; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea kotshubeji; [BRU2]: 64, pl. 38, f. 31-32, pl. 39, f. 1-3; Kolesnichenko, 1999, Atalanta 30 (1/4): 96

Larva on Ligularia thomsonii [BRU2]: 65

• M. k. kotshubeji Peter the Great Mts., Hazreti-Sho, Zaalaisky Mts.

  • Melitaea ala kotshubeji; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 348; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
  • Melitaea chitralensis mod. kotshubeji; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 248, pl. 6, f. 4, 10
  • Melitaea kotshubeji kotshubeji; Kolesnichenko, 1999, Atalanta 30 (1/4): 96, pl. 6, f. 8-11; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. k. bundeli Kolesnichenko, 1999 W.Zaalaisky Mts., S.Alaisky Mts.

  • Melitaea kotshubeji bundeli Kolesnichenko, 1999; Atalanta 30 (1/4): 98, pl. 6, f. 12-13; TL: Alaisky Mts., near Daraut-Kurgan, 3000

  • Melitaea kotshubeji bundeli; [BRU2], 65; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. k. kugarti Kolesnichenko, 1999 northern slopes (E.Alaisky Mts.), Fergansky, Chatkalsky Mts.

  • Melitaea kotshubeji kugarti Kolesnichenko, 1999; Atalanta 30 (1/4): 100, pl. 6, f. 14-17; TL: Fergansky Mts., Kugart river, 2000

  • Melitaea kotshubeji kugarti; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. k. pallidoptera Korb, 2010 Kirghizstan

  • Melitaea kotshubeji pallidoptera Korb, 2010; Eversmannia (21-22): 28; TL: Suusamyr valley (Sary-Kayky mtns)

  • Melitaea kotshubeji pallidoptera; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

• M. e. enarea Garm, Kamara-su river, Peter the Great Mts., Hazreti-Sho, Darvazky Mts., Shugnansky Mts.

  • Melitaea didyma shugnana Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 355, pl. 27, f. 7-10; TL: Chorag
    • = ; Kolesnichenko, 1999, Atalanta 30 (1/4): 106

  • Melitaea chitralensis enarea; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 248, pl. 6, f. 2; Shchetkin, 1981, Ent. Obozr. Tajikistana: 164-165; [LepIndex]; [CoL]
  • Melitaea chitralensis shugnana; Tshikolovets, 1997, The Butterflies of Pamir: 118, pl. 28, f. 7-8, pl. 29, f. 7-8, pl. 43, f. 4; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea didyma enarea; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 333, pl. 1, f. 7, pl. 5, f. 52; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 47
  • Melitaea enarea enarea; Kolesnichenko, 1999, Atalanta 30 (1/4): 106, pl. 7, f. 26-29; [BRU2], 64; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea enarea shugnana; [BRU2], 64
• M. e. ishkashima Sheljuzhko, 1929 Ishkashimsky Mts.

  • Melitaea didyma ishkashima Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 359, pl. 27, f. 13-14; TL: Kishlak Njut
    • = Melitaea enarea shugnana; [BRU2], 64

  • Melitaea didyma ishkashima; Tshikolovets, 1997, The Butterflies of Pamir: pl. 28, f. 13, pl. 29, f. 13, pl. 38, f. 8, pl. 39, f. 8
  • Melitaea enarea ishkashima; Kolesnichenko, 1999, Atalanta 30 (1/4): 108, pl. 7, f. 30
• M. e. permuta Higgins, 1941 Nuratau Mts., Kugitang Mts., Gissarsky Mts., Zeravshansky Mts., Turkenstansky Mts., Karateginsky Mts.

  • Melitaea enarea mod. permuta Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 250, pl. 6, f. 7-8; TL: Samarkand, Hazreth-Sultan

  • Melitaea ala permuta; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
  • Melitaea enarea permuta; Kolesnichenko, 1999, Atalanta 30 (1/4): 109, pl. 7, f. 31
  • Melitaea permuta; [BRU2]: 64, pl. 39, f. 4-6; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. e. gromenkoi Kolesnichenko, 1999 Fergansky Mts.

  • Melitaea enarea gromenkoi Kolesnichenko, 1999; Atalanta 30 (1/4): 110, pl. 7, f. 32-34; TL: Kirghizia, 15km NW from Dzhalal-Abad, Kara-Kianyr Mts., 1200m

  • Melitaea enarea gromenkoi; [BRU2], 64; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

Melitaea sutschana Staudinger, 1892 SE.Siberia, Korea, NE.China. See [maps]

• Melitaea sutschana Staudinger, 1892; in Romanoff, Mém. Lépid. 6: 183; TL: Sutschan-Gebiete
• Melitaea didyma var. sibirica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.)
  • = Melitaea didyma var. didymoides; Staudinger, 1871, Cat. Lepid. eur. Faunegeb. (Edn. 2): 18
• siberica Heyne, [1893]; Palaearkt. Grossschm. 1: 394 (preocc.)
• sutschanala Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 335
• transbaicalica Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 341

• Didymaeformia sutschana; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) sutschana; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea didyma sutschanala; [LepIndex]; [CoL]
• Melitaea didyma transbaicalica; [LepIndex]; [CoL]
• Melitaea sutschana; [BRU2]: 63, pl. 39, f. 16-18; Pazhenkova, Zakharov & Lukhtanov, 2015, ZooKeys 538: 43; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

410x400 (~37Kb) ♀ underside A steppefied meadow on a southern slope of the Polovinnaya Pad' valley few kilometres of the Argun' River left bank 13 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 28th July 1997 (aberrantly large basal black spots), Photo © Oleg Kosterin	410x547 (~53Kb) ♀ upperside A steppefied meadow on the Argun' River left bank terrace 4 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997, Photo © Oleg Kosterin	533x400 (~35Kb) ♂ underside A steppefied southern left slope of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997 (on Leontopodium conglobatum (Turcz.) Hand.-Mazz.), Photo © Oleg Kosterin
565x490 (~34Kb) ♂ upperside A steppefied southern left slope of the Budyumkan River valley 5 km of its confluence with the Argun River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997 (on Leontopodium conglobatum (Turcz.) Hand.-Mazz), Photo © Oleg Kosterin

• M. s. sutschana Transbaikalia - Amur, Ussuri, Korea, NE.China

  • Melitaea sutschana sutschana; [BRU2], 63; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. s. graeseri Gorbunov, 1995 Sakhalin

  • Melitaea sutschana graeseri; [BRU2], 63; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

Larva on Dodartia orientalis [BRU2]: 65

Melitaea lunulata Staudinger, 1901 -. See [maps]

• Melitaea lunulata Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 30; TL: Issyk-Kul

• Didymaeformia lunulata; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) lunulata; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea fergana lunulata; [LepIndex]; [CoL]
• Melitaea lunulata; [BRU2]: 65, pl. 43, f. 16-18; Korb, 2015, Caucasian Ent. Bull. 11 (1): 190; [iNaturalist]

Misidentified/misapplied:

• Melitaea fergana lunulata; Lukhtanov & Lukhtanov, 1994, Herbipoliana 3: pl. 33, f. 5
  • = Melitaea lunulata merke; Lukhtanov, 1999, Atalanta 30 (1/4): 147

• M. l. lunulata Tian-Shan

  • Melitaea lunulata lunulata; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. l. merke Lukhtanov, 1999 W.Tian-Shan

  • Melitaea lunulata merke Lukhtanov, 1999; Atalanta 30 (1/4): 147; TL: Kazakstan, Merke Fl., 2900

  • Melitaea lunulata merke; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

Melitaea fergana Staudinger, 1882 Turkestan. See [maps]

• Melitaea fergana Staudinger, 1882; Berl. ent. Z. 26 (1): 168; TL: Alai-Gebirge [Fergana, Turkestan]
• Melitaea fergana Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 404 (Stgr. i.l.), pl. 14, f. 14-15

• Didymaeformia fergana; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) fergana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• Melitaea fergana; [BRU2]: 66, pl. 42, f. 31-42, pl. 43, f. 1-12; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Melitaea saxatilis var. fergana; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 432, pl. 12, f. 3a-b

• M. f. fergana Alai, Transalai

  • Melitaea fergana fergana; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. f. maracandica Staudinger, 1882 Ghissarsky Mts., Turkestansky Mts. (Pass Kumbel)

  • Melitaea fergana var. maracandica Staudinger, 1882; Berl. ent. Z. 26 (1): 168; TL: Alai (Fergana, Turkestan)
  • ?darvasica Avinoff, 1941
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea fergana splendida Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

  • Didymaeformia macarandica[sic]; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name) [spell.?]
  • Melitaea fergana maracandica; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39; [LepIndex]; [CoL]
  • Melitaea saxatilis var. maracandica; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 433
• M. f. darvasika Higgins, 1941 Darvaz

  • Melitaea fergana darvasika; [BRU2], 66
• M. f. jacobsoni Higgins, 1941 Pamirs

  • Melitaea fergana jacobsoni Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 257
  • Melitaea fergana oxuana Lukhtanov, 1999; Atalanta 30 (1/4): 147; TL: Tadjikistan, E.Pamir, Sarykol, Dunkeidyk, 3750
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
  • Melitaea fergana grumi Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

  • Melitaea fergana grumi; [BRU2], 66
  • Melitaea fergana jacobsoni; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39; [LepIndex]; [CoL]
• M. f. ketmeana Lukhtanov, 1999 S.Kazakstan, NE.Kirghizia

  • Melitaea fergana ketmeana Lukhtanov, 1999; Atalanta 30 (1/4): 145; TL: Kazakstan, Alma-Ata, Ketmen, Kolshat

  • Melitaea fergana ketmeana; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. f. terskeana Lukhtanov, 1999 Kirghizia

  • Melitaea fergana terskeana Lukhtanov, 1999; Atalanta 30 (1/4): 146; TL: Kirghizia, Terskey-Alatoo, Barskoun-Pass, 3400m

  • Melitaea fergana terskeana; [BRU2], 66 (?); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. f. paradoxa Churkin & Tuzov, 2000 N.Tian-Shan

  • Melitaea fergana paradoxa Churkin & Tuzov, 2000; Helios 1: (33-60) [?]

  • Melitaea fergana paradoxa; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39
• M. f. vladislavi Churkin & Tuzov, 2000 Inner Tian-Shan (At-Bashi Mts.)

  • Melitaea fergana vladislavi Churkin & Tuzov, 2000; Helios 1: (33-60) [?]

  • Melitaea fergana vladislavi; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• M. f. khantengri Churkin & Tuzov, 2000 Inner Tian-Shan (Kaindy Mts.)

  • Melitaea fergana khantengri Churkin & Tuzov, 2000; Helios 1: (33-60) [?]

  • Melitaea fergana khantengri; [BRU2], 66; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 39

Melitaea infernalis Grum-Grshimailo, 1891 Tian-Shan, Dzhungarsky Alatau. See [maps]

• Melitaea saxatilis var. infernalis Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 455

• Didymaeformia infernalis; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) infernalis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• Melitaea infernalis; [BRU2]: 67, pl. 43, f. 13-15; [LepIndex] ; [CoL]

• M. i. carmen Churkin & Kolesnichenko, 2002

  • infernalis carmen Churkin & Kolesnichenko, 2002; Helios 3: 148

  • Melitaea infernalis carmen; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40

Melitaea ambrisia Higgins, 1935 Alai, Ghissar. See [maps]

• Melitaea ambrisia Higgins, 1935; Entomologist 68: 112; TL: Alai Mts., Kirgizia

• Didymaeformia ambrisia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) ambrisia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• Melitaea ambrisia; Churkin, 2000, Atalanta 31 (1/2): 114, pl. IV, f. 1-3; [BRU2]: 67, pl. 43, f. 25-30; [LepIndex] ; [CoL]

• M. a. ambrisia Alai

  • Melitaea ambrisia ambrisia; [BRU2], 67; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• M. a. avicaena Churkin, 2000 Turkestansky Mts. (Kumbel Pass)

  • Melitaea ambrisia avicaena Churkin, 2000; Atalanta 31 (1/2): 115, pl. IV, f. 4-6; TL: Uzbekistan, Turkestansky Range, Kumbel Pass, Zaaminsky rs., 3000m

  • Melitaea ambrisia avicaena; [BRU2], 67; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40

Melitaea alraschid Higgins, 1941 -. See [maps]

• Melitaea ambrisia alraschid Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 259; TL: Persia [= error] [?W Gissar, near Sarmarkand]

• Melitaea (Didymaeformia) alrashid[sic, alraschid]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• Melitaea alraschid; Churkin, 2000, Atalanta 31 (1/2): 119, pl. IV, f. 7-9; [BRU2]: 67, pl. 43, f. 19-24; [LEPIFORUM]
• Melitaea ambrisia alraschid; [LepIndex]; [CoL]

• M. a. alraschid Ghissar

  • Melitaea alraschid alraschid; [BRU2], 67
• M. a. garun Churkin, 2000 Matcha

  • Melitaea alraschid garun; [BRU2], 67
• M. a. irinae Churkin, Tremasov & Kolesnichenko, 2012 Tadjikistan (Peter the Great Range)

  • Melitaea alraschid irinae Churkin, Tremasov & Kolesnichenko, 2012; Atalanta 43: 118; TL: Tadjikistan, Peter the Great Mts, 23km SE Tadzhikobad, Ganishou v., 3400-3500m
    • = Melitaea fergana maracandica; Korb & Bolshakov, 2016, Zootaxa 4160 (1): (1-234)

Melitaea shandura Evans, 1924 Afghanistan, Pamirs, Pakistan, NW.India. See [maps]

• Melitaea shandura Evans, 1924; J. Bombay nat. Hist. Soc. 30 (1): 90; TL: Shandur, Chitral [NW.India]
• Melitaea saxatilis shandura Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 583 (Christoph) (nom. nud.)

• Didymaeformia shandura; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) shandura; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• Melitaea shandura; [BRU2]: 68, pl. 43, f. 31-33; [LepIndex] ; [CoL]

• M. s. shandura

  • Melitaea shandura shandura; Sakai, 1978, Atalanta 9 (1): 109
• M. s. pavlitzkajana Sheljuzhko, 1943 W.Pamirs

  • Melitaea pavlitzkajana Sheljuzhko, 1943; Mitt. Münchn. ent. Ges. 33: 75
  • Melitaea shandura apsara Wyatt, 1961; J. Lep. Soc. 15 (1): 6; TL: above Kars Pass, Shiva Mts, N. Badakhshan, 12500-14000ft, Afghanistan

  • Melitaea pavlitzkajana; [LepIndex] ; [CoL]
  • Melitaea shandura apsara; [LepIndex]; [CoL]
  • Melitaea shandura pavlitzkajana; [BRU2], 68; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40

• M. a. danae Churkin & Kolesnichenko, 2005

  • Melitaea athene danae; Yakovlev, 2012, Nota lepid. 35 (1): 89

Melitaea trivia (Denis & Schiffermüller, 1775) -. See [maps] Baluchi Fritillary (_robertsi_) Desert Fritillary Lesser Spotted Fritillary Braunlicher Scheckenfalter

• Papilio trivia Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 179; TL: Vienna
  • = Melitaea fascelis; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
• Papilio phoebe Esper, 1782; Die Schmett. Th. I, Bd. 2 (8): 172, pl. 88, f. 5-6
• Papilio antigonus Herbst, 1800; Naturs. Schmett. 10: pl. 278, f. 5-8
• Melitaea robertsi Butler, 1880; Proc. zool. Soc. Lond. 1880 (3): 406, pl. 39, f. 2
  • = ; [Wahlberg]
• Melitaea didyma ab. interrupta Skala, 1907; Ent. Zs. 20 (42): 311
• Melitaea didyma ab. radiata Skala, 1907; Ent. Zs. 20 (42): 311, pl. 4, f. 8
• Melitaea didyma ab. hoffmanni Skala, 1908; Ent. Zs. 21: 268 (repl. Melitaea didyma ab. radiata Skala, 1907)
• Melitaea trivia ab. tenuisignata Skala, 1912; Ver. naturf. Ver. Brünn 50: 109
• Melitaea trivia ab. latemarginata Grund, 1913; Int. ent. Zs. 7 (19): 127
• Melitaea trivia ab. ornata Grund, 1913; Int. ent. Zs. 7 (19): 127
• Melitaea trivia ab. variegata Grund, 1913; Int. ent. Zs. 7 (19): 127
• Melitaea trivia ab. melanina Grund, 1913; Int. ent. Zs. 7 (19): 127
• Melitaea trivia catapelioides Stauder, 1918; Zs. wiss. Insektenbiol. 14 (3/4): 57
• Melitaea trivia f. striata Gusic, 1922; Neue Beitr. syst. Ins. 2 (10): 96
• Melitaea trivia f. nigrofasciata Gusic, 1922; Neue Beitr. syst. Ins. 2 (10): 96
• Melitaea trivia ab. ardens Erfurth, 1923; Zs. Öst. EntVer. 8: 56
• equivalens de Sagarra, 1924; Trab. mus. Ciénc. nat. Barcelona 11 (1): 15
• Melitaea trivia ignasiti de Sagarra, 1926; Butll. Inst. catal. Hist. nat. (2) 6 (6-7): 131; TL: Portugal
  • = ; [FE]
• trivia tigris Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 286
• Melitaea trivia f. fasciata Eisner, 1942; Zool. Meded. Leiden 24 (4): 118; TL: Hungary
• Melitaea trivia f. flavescens Eisner, 1942; Zool. Meded. Leiden 24 (4): 118; TL: Krain; Edessa (Macedonien)
• Melitaea trivia blötei [= blotei] Eisner, 1942; Zool. Meded. Leiden 24 (4): 119
• Melitaea trivia gen. brussaensis Eisner, 1942; Zool. Meded. Leiden 24 (4): 119; TL: Karacabey, Brussa
• Melitaea trivia gen. vernalis Eisner, 1942; Zool. Meded. Leiden 24 (4): 120; TL: Syria
• quercii Rocca, 1946; Boll. Soc. ent. ital. 76: 36
• catamesoides Verity, 1950; Le Farfalle diurn. d'Italia 4: 97
• catananoides Verity, 1950; Le Farfalle diurn. d'Italia 4: 97
• microignasita Verity, 1950; Le Farfalle diurn. d'Italia 4: 97
• microsyriaca Verity, 1950; Le Farfalle diurn. d'Italia 4: 97
• Melitaea pusilla Heydemann, [1955]; Zs. Wien. Ent. Ges. 39 (12): 420, pl. 36, f. 11; TL: Sarobi
  • = Melitaea catapelia; Shchetkin, 1984, Zool. Zh. 64: (1822-1827)

• Didymaeformia trivia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea (Didymaeformia) robertsi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• Melitaea (Didymaeformia) trivia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
• Melitaea ignasiti; [iNaturalist] ; [LEPIFORUM]
• Melitaea robertsi; [iNaturalist] ; [LEPIFORUM]
• Melitaea trivia blotei; [LepIndex]; [CoL]
• Melitaea trivia catamesoides; [LepIndex]; [CoL]
• Melitaea trivia catananoides; [LepIndex]; [CoL]
• Melitaea trivia ignasiti; [H&R], 118
• Melitaea trivia nigrofasciata; [LepIndex]; [CoL]
• Melitaea trivia; Fruhstorfer, 1919, Archiv Naturg. 83 A (6): 12; [H&R], 118; [BOW]: pl. 3, f. 5; [EBW]; [BRU2]: 62, pl. 40, f. 1-36; [Otakar Kudrna]; [FE]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [Mario Maier]

Larva on Verbascum thapsus [H&R]

• M. t. trivia

  • Melitaea trivia trivia; [H&R], 118; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40; [iNaturalist]
• M. t. fascelis (Esper, 1783) S.Europe

  • Papilio fascelis Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 171, pl. 88, f. 3-4
    • = Melitaea trivia; [FE]
  • Papilio iphigenia Esper, 1782; Die Schmett. Th. I, Bd. 2 (5): 128, pl. 77, f. 1-2 (preocc.)
    • = Melitaea fascelis fascelis; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • cleo Latreille, 1803
    • = Melitaea fascelis fascelis; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • minor Ksienschopolski, 1911
  • striata Ksienschopolski, 1911; Trudy obshch.Izsl. Volyni Zitomir 8: 38, pl. 1, f. 6
  • Melitaea trivia paula Obraztsov, 1936; Festschrift für Dr. Embrik Strand 1: 640; TL: Kiev

  • Melitaea fascelis fascelis; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea fascelis; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45; [iNaturalist]
  • Melitaea trivia fascelis; [BRU2], 62
  • Melitaea trivia paula; [LepIndex]; [CoL]
  • Papilio fascelis; Fabricius, 1787, Mantissa Insectorum 2: 58

  Larva on Scrophularia , Eremostachys [BRU2]: 62

• M. t. nana Staudinger, 1871 Turkey, Iraq, Iran

  • Melitaea trivia var. nana Staudinger, 1871; Cat. Lepid. eur. Faunegeb. (Edn. 2): 18
  • Melitaea didyma aabaca Fruhstorfer, 1917; Soc. ent. 32 (5): 19; TL: Spain, Castilia
    • = Melitaea fascelis nana; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • wyatti Gross & Ebert, 1975
    • = Melitaea fascelis nana; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45

  • Melitaea fascelis nana; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea trivia nana; [BRU2], 62; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40; [LepIndex]; [CoL] ; [iNaturalist]
• M. t. catapelia Staudinger, 1886 Ghissar, Alai, Tian-Shan, Pamirs

  • Melitaea trivia var. catapelia Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 231
  • Melitaea catapelia bactriana Shchetkin, 1984; Zool. Zh. 64: (1822-1827)
    • = ; [BRU2], 62
    • = Melitaea robertsi catapelia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
  • Melitaea trivia nativa Tuzov, 2000; [BRU2], 62 (repl.)
    • = Melitaea robertsi catapelia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
  • Melitaea catapelia shugnana Shchetkin, 1984; Zool. Zh. 64: (1822-1827) (preocc. Melitaea didyma shugnana Sheljuzhko, 1929)

  • Melitaea robertsi catapelia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
  • Melitaea trivia catapelia; [BRU2], 62
  • Melitaea trivia var. catapelia; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 429
• M. t. pseudodidyma Rebel, 1905 Turkey

  • Melitaea trivia var. pseudodidyma Rebel, 1905; Ann. nat. Mus. Wien 20: 194; TL: Brussa

  • Melitaea fascelis pseudodidyma; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea trivia pseudodidyma; [LepIndex]; [CoL]
• M. t. syriaca Rebel, 1905 Lebanon, Jordan, Israel

  • Melitaea trivia var. syriaca Rebel, 1905; Ann. nat. Mus. Wien 20: 194; TL: Syria

  • Melitaea fascelis syriaca; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 45
  • Melitaea syriaca; [iNaturalist] ; [LEPIFORUM]
• M. t. mercea Evans, 1912

  • Melitaea trivia mercea Evans, 1912; J. Bombay nat. Hist. Soc. 21 (3): 1008

  • Melitaea trivia mercea; [LepIndex]; [CoL]
• M. t. lathon Fruhstorfer, 1917 Bosnia

  • Melitaea trivia lathon Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 13; TL: Velebit, Bosnia
  • infrajuncta Verity, 1950; Le Farfalle diurn. d'Italia 4: 98

  • Melitaea trivia lathon; [LepIndex]; [CoL]
• M. t. stemmleri Collier, 1933 Macedonia

  • trivia stemmleri Collier, 1933; Ent. Rundsch. 50: 55

  • Melitaea trivia stemmleri; [LepIndex]; [CoL]
• M. t. salamancaensis Eisner, 1942 Spain

  • Melitaea trivia salamancaensis Eisner, 1942; Zool. Meded. Leiden 24 (4): 119; TL: Peña del Francia, Salamanca

  • Melitaea trivia salamancaensis; [LepIndex]; [CoL]
• M. t. djugjurensis Eisner, 1942 Amur

  • Melitaea trivia djugjurensis Eisner, 1942; Zool. Meded. Leiden 24 (4): 120; TL: Djugjur, Amur

  • Melitaea trivia djugjurensis; [LepIndex]; [CoL]
• M. t. chorasana Shchetkin, 1984 S.Ghissar

  • Melitaea catapelia chorasana Shchetkin, 1984; Zool. Zh. 64: (1822-1827)

  • Melitaea robertsi chorasana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
  • Melitaea trivia chorasana; [BRU2], 62
• M. t. petri Shchetkin, 1984 Darvaz

  • Melitaea catapelia petri Shchetkin, 1984; Zool. Zh. 64: (1822-1827)

  • Melitaea trivia petri; [BRU2], 62

  Larva on Scrophularia integrifolia, S. kiriloviana Shchetkin, 1984, Zool. Zh. 64

• M. t. uvarovi Gorbunov, 1995 Kazakhstan, SW.Siberia, Tarbagatai, Saur, Dzhungarsky Alatau

  • Melitaea robertsi uvarovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40 (?); [SZMN]
  • Melitaea trivia uvarovi; [BRU2], 62
• M. t. gulasinaria Korb, 2005 Tuva

  • Melitaea fascelis singularia Korshunov, 1995
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40 (nom. nud.)

  • Melitaea trivia gulasinaria; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 40
  • Melitaea trivia singularia; [BRU2], 62; Yakovlev, 2012, Nota lepid. 35 (1): 90; [SZMN]

Melitaea yuenty Oberthür, 1886 W.China. See [maps]

• Melitaea yuenty Oberthür, 1886; Étud. d'Ent. 11: 17, pl. 2, f. 13; TL: Tatsienlu

• Didymaeformia yuenty; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
• Melitaea yuenty; [BOW]: pl. 196, f. 9; [MRS], 587; [LepIndex] ; [CoL] ; [iNaturalist]

• M. y. yuenty

  • Melitaea yuenty yuenty; [Yutaka Inayoshi]
• M. y. batangensis Belter, 1944 Tibet

  • Melitaea yuenty batangensis Belter, 1944; Ent. Zs. 57: 173; TL: Batang

  • Melitaea yuenty batangensis; Huang, 2001; [LepIndex]; [CoL]

Melitaea romanovi Grum-Grshimailo, 1891 N.China, Mongolia, Transbaikalia. See [maps]

• Melitaea romanovi Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 454; TL: [Gansu, China]

• Melitaea (Melitaea) romanovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42
• Melitaea romanovi; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [MRS], 587; [BRU2]: 65, pl. 44, f. 16-18; [LepIndex] ; [CoL] ; [iNaturalist]

• M. r. romanovi N.China

  • Melitaea romanovi romanovi; [BRU2], 65
• M. r. puella Higgins, 1941 Transbaikalia

  • Melitaea romanovi puella Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 274

  • Melitaea romanovi puella; [Oleg Kosterin]; [BRU2], 65; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42; [SZMN]

  800x600 (~103Kb) ♀ underside The southern slope, covered with a short-grass steppe, of the hill Chikhalan on the northern bank of a brackish Lake Zun-Torei, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 12th July 1996, Photo © Oleg Kosterin

  600x467 (~24Kb) ♀ A short-grass steppe, on the northern bank of a brackish Lake Zun-Torei at the foot of the hill Chikhalan, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 13th July 1996 (captured by a spider), Photo © Oleg Kosterin

• M. r. shanshiensis (Murayama, 1955) N.China (Shansi)

  • Euphidryas[sic] romanovi shanshiensis Murayama, 1955; Tyô to Ga 6 (1): 1, f. 2; TL: Tai-yuan, Shansi, North China

  • Melitaea romanovi shanshiensis; [LepIndex]; [CoL]

Melitaea arduinna (Esper, 1783) SEEU, S.Russia, Asia Minor, Iran, C.Asia. See [maps] Freyer's Fritillary

• Papilio arduinna Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 169, pl. 87, f. 4; TL: [Kamyshin, Volgograd Region, Russia]

• Melitaea (Melitaea) arduinna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42
• Melitaea arduinna; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [H&R], 114; [BOW]: pl. 2, f. 28 (text only); [BRU2]: 72, pl. 45, f. 1-9; [Otakar Kudrna]; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46; [FE]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]

1128x901 (~303Kb) ♂ upperside Iran, Ostan-e Markazi, Shakhrestan-e Shazand, a brook valley downstream Baneh village. 19th May 2017, Photo © Oleg Kosterin

975x882 (~253Kb) ♂ Iran, Ostan-e Markazi, Shakhrestan-e Shazand, a brook valley downstream Baneh village. 19th May 2017, Photo © Oleg Kosterin

856x910 (~274Kb) ♂ upperside Iran, Ostan-e Markazi, Shakhrestan-e Shazand, a brook valley downstream Baneh village. 19th May 2017, Photo © Oleg Kosterin

Larva on Centaurea behen, (Transcaucasia) [BRU2]: 72

• M. a. arduinna SEEU, SW.Siberia, Altai, Alai, N.Tian-Shan, Dzhungarsky Alatau, Tarbagatai, Saur

  • Melitaea rhodopensis Freyer, [1836]; Neuere Beitr. Schmett. 3: 3, pl. 193, f. 1; TL: ?Macedonia "europäischen Türkei"
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
    • = Melitaea arduinna; [FE]
  • arduinna uralensis Eversmann, 1844; [??]
    • = Melitaea arduinna; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
  • ?uralensis Eversmann, 1851; Ent. Russ. 5: 77, pl. 10, f. 1-4

  • Melitaea arduinna arduinna; [BRU2], 72; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
  • Melitaea arduinna rhodopensis; [H&R], 114; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42
  • Melitaea arduinna uralensis; [BRU2], 72
  • Melitaea arduinna var.? rhodopensis; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 427
  • rhodopensis; Abadjiev, 2002, Neue Ent. Nachr. 53: 45
• M. a. evanescens Staudinger, 1886 Kopet-Dagh, Ghissar-Darvaz, W.Pamirs

  • Melitaea arduinna var. evanescens Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 230
  • Melitaea arduinna ab. fulminans Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 230
    • = ; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46

  • Melitaea arduinna evanescens; [BRU2], 72; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 42; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46
  • Melitaea arduinna ab. fulminans; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 427
  • Melitaea arduinna var. evanescens; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 427
• M. a. kocaki Wagener & Gross, 1976 Transcaucasus, Turkey

  • Melitaea arduinna kocaki Wagener & Gross, 1976; Ent. Zs. 86 (20): 229

  • Melitaea arduinna kocaki; [BRU2], 72; Kemal & Koçak, 2011, Priamus 25 (Suppl.): 46; [LepIndex]; [CoL] ; [iNaturalist]